Evolution typically happens over long time periods, with organisms being selected based on their environments. But climate change and human factors can also increase the rate of evolution. In this module we show how organisms evolve in terms of their environment, their competitors, and human influences. The module will begin by exploring the different ways in which interacting organisms can influence each other's evolution, and how co-evolution manifest itself in different types of interactions. Then we will focus on the interaction between humans, animals and plants and how domestication has given rise to the society we see today. An understanding of evolutionary processes, using domestication as a model system, will be investigated. We will then use example of recent evolution to understand how selection is manifest at the genomic level and how species evolve and adapt.